Wolf Dens in Ghost of Yotei are the only way to obtain the skills point you can use to upgrade Atsu's wolf companion's abilities. To complete these, which usually involves freeing wolves from traps, you'll first have to find the dens in the first place. They appear as small caves with a tree on top (as shown below) and you'll get clues from talking to people, and hearing wolf howls when you're out exploring, but it's a big world so I can help point you in the right direction more quickly.
Below is the location of all of the Ghost of Yotei Wolf Dens, broken down by region. Listen out for those wolf howls when you get near the general area, or speak to anyone you pass to help narrow things down. Unlike the Ghost of Yotei Altars of Reflection that will let you upgrade all your Ghost of Yotei skills , wolf skills can only be unlocked by completing the dens I've listed below.
Ghost of Yotei Grasslands Wolf Den Locations
- Great Mountain Wolf Den
- Green Hill Wolf Den
Ghost of Yotei Tokachi Range Wolf Den Locations
- Howling Fields Wolf Den
- Hunter's Watch Wolf Den
Ghost of Yotei Ishikari Plain Wolf Den Locations
- Old Wound Wolf Den
- Dying Fire Wolf Den
Ghost of Yotei Nayoro Wilds Wolf Den Locations
- White Scars Wolf Den
Ghost of Yotei Teshio Ridge Wolf Den Locations
- Biting Wind Wolf Den
- Cold Gale Wolf Den
Ghost of Yotei Oshima Coast Wolf Den Locations
- Waterfall Wolf Den
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